Monilophytes of Maine
(Kingdom Plantae)


a project of
V. F. Thomas Co. - 167 Thorne Mountain Road - Canton, Maine  04221
info@vfthomas.com


(updated 25 December 2019)


Below is a working checklist of Maine’s monilophytes (hybrids excluded). Taxonomy and nomenclature follow Flora Novae Angliae by Arthur Haines. Please send corrections and updates to info@vfthomas.com or mail them to the address at the top of this page. Photographs are welcome. Thank you.


Family Genus Specific Epithet authority Sub-specific Epithet authority common name Sources (full citations given after this table)
Aspleniaceae Asplenium platyneuron (L.) Britton mountain spleenwort
Aspleniaceae Asplenium rhizophyllum L. walking spleenwort
Aspleniaceae Asplenium trichomanes L. ssp. trichomanes   maidenhair spleenwort Greene et al. (2005)
Aspleniaceae Asplenium viride Huds. green spleenwort
Blechnaceae Woodwardia aereolata (L.) T. Moore     netted chain fern
Blechnaceae Woodwardia virginica (L.) Smith     Virginia chain fern Greene et al. (2005)
Dennstaedtiaceae Dennstaedtia punctilobula (Michx.) T. Moore     eastern hay-scented fern Champlainer (TFV); Greene et al. (2005)
Dennstaedtiaceae Pteridium aquilinum (L.) Kuhn ssp. latiusculum (Desv.) Hultén bracken fern Champlain Project (2007); Greene et al. (2005)
Dryopteridaceae Dryopteris campyloptera (Kunze) Clarkson     mountain wood fern Champlain Project (2007); Greene et al. (2005)
Dryopteridaceae Dryopteris carthusiana (Vill.) H. P. Fuchs     spinulose wood fern Greene et al. (2005)
Dryopteridaceae Dryopteris clintoniana (D. C. Eat.) Dowell     Clinton’s wood fern
Dryopteridaceae Dryopteris cristata (L.) Gray     crested wood fern Greene et al. (2005)
Dryopteridaceae Dryopteris filix-mas (L.) Schott ssp. brittonii Fras.-Jenk. & Widen Clinton’s wood fern
Dryopteridaceae Dryopteris fragrans (L.) Schott     fragrant wood fern
Dryopteridaceae Dryopteris goldiana (Hook. ex Goldie) Gray     Goldie’s wood fern
Dryopteridaceae Dryopteris intermedia (Muhl. ex Willd.) Gray     evergreen wood fern Champlain Project (2007); Greene et al. (2005)
Dryopteridaceae Dryopteris marginalis (L.) Gray     marginal wood fern Champlain Project (2007); Greene et al. (2005)
Dryopteridaceae Polystichum acrostichoides (Michx.) Schott     Christmas fern Champlain Project (2007); Greene et al. (2005)
Dryopteridaceae Polystichum braunii (Spenner) Fée     Braun’s holly fern
Equisetaceae Equisetum arvense L.     field horsetail Champlainers (TFV, EMW); Greene et al. (2005)
Equisetaceae Equisetum fluviatile L.     river horsetail Champlainer (TFV); Greene et al. (2005)
Equisetaceae Equisetum hyemale L. ssp. affine (Engelm.) Calder & Taylor tall scouring-rush
Equisetaceae Equisetum palustre L. marsh horsetail
Equisetaceae Equisetum pratense Ehrh. meadow horsetail
Equisetaceae Equisetum scirpoides Michx. dwarf scouring-rus
Equisetaceae Equisetum sylvaticum L.     wood horsetail Champlainers (AH, TFV); Greene et al. (2005)
Equisetaceae Equisetum variegatum Schleicher ex F. Weber & D. Mohr ssp. variegatum variegated scouring-rush
Marsileaceae Marsilea quadrifolia L. European water-clover
Onocleaceae Matteuccia struthiopteris (L) Todaro ssp. pensylvanica (Willd.) A. & D. Love fiddlehead fern
Onocleaceae Onoclea sensibilis L.     sensitive fern Champlain Project (2007); Greene et al. (2005)
Ophioglossaceae Botrychium angustisegmentum (Pease & Moore) Fern.     narrow triangle moonwort
Ophioglossaceae Botrychium dissectum Spreng.     dissected grapefern
Ophioglossaceae Botrychium lunaria (L.) Sw.     common moonwort
Ophioglossaceae Botrychium matricariifolium (Doll) A. Braun ex Koch     dissected grapefern
Ophioglossaceae Botrychium minganense Victorin     Mingan moonwort
Ophioglossaceae Botrychium multifidum (Gmel.) Rupr.     leathery grapefern
Ophioglossaceae Botrychium oneidense (Gilbert) House     blunt-lobed grapefern
Ophioglossaceae Botrychium pallidum W. H. Wagner     pale moonwort
Ophioglossaceae Botrychium simplex E. Hitchc.     least moonwort
Ophioglossaceae Botrychium tenebrosum A. A. Eat.     swamp moonwort
Ophioglossaceae Botrychium virginianum (L.) Sw.     rattlesnake fern Greene et al. (2005)
Ophioglossaceae Ophioglossum pusillum Raf.     northern adder’s-tongue fern Greene et al. (2005)
Osmundaceae Osmunda claytoniana L.     interrupted fern Champlain Project (2007); Greene et al. (2005)
Osmundaceae Osmunda regalis L. var. spectabilis (Willd.) Gray royal fern Champlainer (TFV); Greene et al. (2005)
Osmundaceae Osmundastrum cinnamomeum (L.) C. Presl     cinnamon fern Champlain Project (2007); Greene et al. (2005) as Osmunda cinnamomea
Polypodiaceae Polypodium appalachianum Haufler & Windham     Appalachian polypody Greene et al. (2005)
Polypodiaceae Polypodium virginianum L.     rock polypody Greene et al. (2005)
Pteridaceae Adiantum aleuticum (Rupr.) Paris     western maidenhair fern
Pteridaceae Adiantum pedatum L.     northern maidenhair fern Greene et al. (2005)
Pteridaceae Adiantum viridimontanum Paris     Green Mountain maidenhair fern
Pteridaceae Cryptogramma stelleri (Gmel.) Prantl     slender rock-brake
Thelypteridaceae Parathelypteris noveboracensis (L.) Ching     New York fern Champlain Project (2007); Greene et al. (2005) as Thelypteris noveboracensis
Thelypteridaceae Parathelypteris simulata (Davenport) Holttum     Massachusetts fern
Thelypteridaceae Phegopteris connectilis (Michx.) Watt     long beech fern Champlain Project (2007); Greene et al. (2005)
Thelypteridaceae Phegopteris hexagonoptera (Michx.) Fée     broad beech fern
Thelypteridaceae Thelypteris palustris Schott var. pubescens (Lawson) Fern. marsh fern Greene et al. (2005)
Woodsiaceae Athyrium angustum (Willd.) C. Presl     narrow lady fern Champlainer (TFV); Greene et al. (2005) as Athyrium filix-femina var. angustum
Woodsiaceae Cystopteris bulbifera (L.) Bernh.     bulbil fragile fern
Woodsiaceae Cystopteris fragilis (L.) Bernh.     fragile fern Greene et al. (2005)
Woodsiaceae Cystopteris tenuis (Michx.) Desv.     Mackay’s fragile fern
Woodsiaceae Deparia acrostichoides (Sw.) M. Kato     silvery false spleenwort Greene et al. (2005)
Woodsiaceae Gymnocarpium dryopteris (L.) Newman     northern oak fern Champlainer (TFV); Greene et al. (2005)
Woodsiaceae Woodsia alpina (Bolton) S. F. Gray     northern rusty cliff fern
Woodsiaceae Woodsia glabella R. Br. ex Richards.     smooth cliff fern
Woodsiaceae Woodsia ilvensis (L.) R. Br.     rusty cliff fern Champlainer (TFV); Greene et al. (2005)
Woodsiaceae Woodsia obtusa (Spreng.) Torr. ssp. obtusa   blunt-lobed cliff fern


Sources:
   Champlain Project (2007) = Champlain Project fern walk on 27 August 2007 along Valley Trail (Mount Desert Island trail 116).
   Champlainers:
      AH = Arthur Haines
      TFV = Thomas F. Vining
      EMW = Emma Walsh
   Greene, Craig W., Linda L. Gregory, Glen H. Mittelhauser, Sally C. Rooney, and Jill E. Weber. 2005. Vascular Flora of the Acadia National Park Region. Rhodora 107(930):117-185.